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Suitable alternative to Quicktime RLE for alpha-containing non-cartoon videos?

ProRes 4444 is the go-to for compressed encoding if you want alpha, and you’re doing the encoding on a Mac. It’s not exactly lightweight in terms of file size, but it’s optimized for playback on Mac ...
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Widely compatible video format with lossless audio

According to Mozilla Developers MPEG-4 containers can include FLAC audio. Ditto for OGG containers.
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